Michael S. Hedrick


Department of Biological Sciences

(510) 885-7443
SC-S 351
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Photo of Michael Hedrick

I am a comparative physiologist with primary interests in respiratory, cardiovascular and exercise physiology in ectothermic vertebrates. Recent projects include regulation of lymphatics and baroreflexes in anuran amphibians, regulation of respiratory rhythm generation during development in amphibians, and meta-analyses of maximal O2 and CO2 flux rates in vertebrates.

Professional Focus

Professional Appointments and Positions

2016-present   Director, Semester Conversion Initiative, CSU-East Bay

2005-present   Professor, Dept. of Biological Sciences, CSU-East Bay

2011-2014       Professor, Dept. of Biological Sciences, Univ. of North Texas, Denton, TX

2008-2011       Chair, Dept. of Biological Sciences, CSU-East Bay

2000-2005       Associate Professor, Dept. of Biological Sciences, CSU-East Bay (name change 2005)

1994-2000       Assistant Professor, Dept. of Biological Sciences, CSU-Hayward, Hayward, CA

1996                Visiting Assistant Professor, University of Wisconsin School of Veterinary Medicine

1994                Research Associate, Dept. of Comparative Biosciences, Univ. of Wisconsin

1991-93           Postdoctoral Fellow, Dept. of Comparative Biosciences, School of Veterinary Medicine.  University of Wisconsin, Madison, WI


  • PhD., 1991. Zoology. University of British Columbia
  • M.S., 1985. Biology. Portland State University
  • B.S., 1980. Biology. Lewis and Clark College


Not teaching this quarter.


Recent Publications

Peer-Reviewed Journal Publications

Hillman, S.S. and Hedrick, M.S. (2017). A metabolic hypothesis for the evolution of temperature effects on the aterial PCO2 and pH of vertebrate ectotherms. Journal of Experimental Biology. In Review.

Enok, S., Leite, G., Leite, C., Gesser, H. Hedrick, M.S. and Wang, T. (2016). Improved cardiac filling facilitates the postprandial elevation of stroke volume in Python regius. Journal of Experimental Biology. 219: 3009-3018. 

Johnson, S.M., Hedrick, M.S., Krause, B.M., Nilles, J.P. and Chapman, M.A. (2016). Respiratory neuron characterization reveals intrinsic bursting properties in isolated adult turtle brainstems (Trachemys scripta). Respiratory Physiology and Neurobiology. 224: 52-61.

Hedrick, M.S., Hancock, T.V. and Hillman, S.S. (2015). Metabolism at the max: how vertebrate organisms respond to physical activity. Comprehensive Physiology. 5: 1677-1703.

Hillman, S.S. and Hedrick, M.S. (2015). A meta-analysis of in vivo vertebrate cardiac performance: implications for cardiovascular support in the evolution of endothermy. Journal of Experimental Biology. 218: 1143-1150. Featured 'Inside JEB' article.

Hedrick, M.S., McNew, K.A. and Crossley II, D.A. (2015). Baroreflex function in anurans from different environments. Comparative Biochemistry and Physiology Part A. 179: 144-148.

Hillman, S.S., Drewes, R.C., Hedrick, M.S. and Hancock, T.V. (2014). Physiological vagility and its relationship to dispersal and neutral genetic heterogeneity in vertebrates. Journal of Experimental Biology 217: 3356-3364.

Hedrick, M.S., Hansen, K., Wang, T., Lauridsen, H., Thygesen, J. and Pedersen, M. (2014). Visualising lymph movement in anuran amphibians with computed tomography. Journal of Experimental Biology 217: 2990-2993. Featured ‘Inside JEB’ article.

Hillman, S.S., Hedrick, M.S. and Kohl, Z.F. (2014). Net cardiac shunts in anuran amphibians: physiology or physics? Journal of Experimental Biology 217: 2844-2847.

Marschand, R., Wilson, J., Burleson, M.L., Crossley, D.A. and Hedrick, M.S. (2014). Effects of prolonged lung inflation or deflation on pulmonary stretch receptor discharge in the alligator (Alligator mississippiensis). Respiratory Physiology and Neurobiology 200: 25-32.

Buckley, B.A., Hedrick, M.S. and Hillman, S.S. (2014). Cardiovascular oxygen transport limitations to thermal niche expansion and environmental PO2 in Antarctic notothenioid fishes. Physiological and Biochemical Zoology 87: 499-506.

Withers, P.C., Hedrick, M.S., Drewes, R.C. and Hillman, S.S. (2014). Pulmonary compliance and lung volume are related to terrestriality in anuran amphibians. Physiological and Biochemical Zoology  87: 374-383.

Turesson, J., Hedrick, M.S., Sundin, L. and Burleson, M.L. (2014). Brainstem mechanisms controlling cardiovascular reflexes in channel catfish. Comparative Biochemistry and Physiology Part. A. 170: 1-5.

Hillman, S.S., Drewes, R.C., Hedrick, M.S. and Hancock, T.V. (2014).  Physiological vagility: Correlations with dispersal and population genetic structure of amphibians. Physiological and Biochemical Zoology 87: 105-112.

Burggren, W.W., Christoffels, V.M., Crossley, D.A., Enok, S., Farrell, A.P., Hedrick, M.S., Hicks, J.W., Jensen, B., Moorman, A.F.M., Mueller, C., Skovgaard, N., Taylor, E.W. and Wang, T. (2014). Comparative cardiovascular physiology: Future trends, opportunities and challenges. Acta Physiologica 210: 257-276.

Hedrick, M.S., Hillman, S.S., Drewes, R.C. and Withers, P.C. (2013). Lymphatic regulation in nonmammalian vertebrates. Journal of Applied Physiology 115: 297-308.

Kohl, Z.F., Hedrick, M.S. and Hillman, S.S. (2013). Separating the contributions of vascular anatomy and blood viscosity to peripheral resistance and the physiological implications of interspecific resistance variation in amphibians. Journal of Comparative Physiology B. 183: 921-932.

Drewes, R.C., Hillman, S.S., Hedrick, M.S. and Withers, P.C. (2013). Evolutionary implications of the distribution and variation of the skeletal muscles of the anuran lymphatic system. Zoomorphology. 132: 339-349.

Klingler, M.J. and Hedrick, M.S. (2013). Evidence for rhombomeric organization of multiple respiratory oscillators during development in the bullfrog brainstem. Respiratory Physiology and Neurobiology 186: 7-15.

Hillman, S.S., Hancock, T.V. and Hedrick, M.S. (2013). A comparative meta-analysis of maximal aerobic metabolism of vertebrates: Implications for respiratory and cardiovascular limits to gas exchange. Journal of Comparative Physiology B. 183: 167-179.

Book Chapters

Hedrick, M.S. and Katz, S.L. Control of breathing in primitive fishes. In: Phylogeny, Anatomy and Physiology of Ancient Fishes. (edited by Giacomo Zaccone, Konrad Dabrowski, Michael S. Hedrick, J. M.O. Fernandes and Jose M. Icardo). CRC Press: Boca Raton, FL, 2016, pp. 179-200.


Zaccone, G., Dabrowski, K., Hedrick, M.S., Fernandes, J.M.O and Icardo, J. (editors). Phylogeny, Anatomy and Physiology of Ancient Fishes. CRC Press: Boca Raton, FL, 2016, 297 pp.

Other Publications

Hillman, S.S., Hedrick, M.S. and Kohl, Z.F. (2017). Commentary on: "Vascular distensiblities have minor effects on intracardiac shunt patterns in reptiles" Zoology. 122: 52-54.

Hedrick, M.S. and Hillman, S.S. (2016). What drove the evolution of endothermy? JEB Classics: Endothermy and Activity in Vertebrates, by A.F. Bennett and J.A. Ruben. Journal of Experimental Biology 219: 300-301.


Recent Invited Seminars

June, 2016      University of Aarhus, Denmark, Dept. of Zoophysiology

May, 2013       University of Aarhus, Denmark, Dept. of Zoophysiology

Mar., 2013       Queen’s University, Canada, Dept. of Biology

Jan., 2013       Texas Woman’s University, Dept. of Biology

Oct., 2012       Wright State University, Dept. of Biological Sciences

Sep., 2012       Saint Louis University, Dept. of Biology

Mar., 2012       University of Oklahoma, Dept. of Zoology

Nov., 2011       University of Toronto, Scarborough, Dept. of Biological Sciences

Recent Conference Presentations/Abstracts

Hedrick, M.S. and Hillman, S.S. (2018). A metabolic hypothesis for the evolution of temperature effects on the arterial PCO2 and pH of ectothermic vertebrates. Soc. Int. Comp. Biol. San Francisco, CA.

Hedrick, M.S., Streicher, J.W., Crossley II, D.A. and Fujita, M.K. (2015). Maximal metabolic rates of the Barking frog (Craugastor augusti). Exp. Biol. Boston, MA.

Hedrick, M.S. and Crossley II, D.A. (2014). Baroreflex characteristics of anuran amphibians from different environments. Am. Physiol. Soc. Intersociety Meeting. San Diego, CA.

Hedrick, M.S., Hansen, K., Wang, T., Thygesen, J. and Pedersen, M. (2014). Visualizing lymph movement in anuran amphibians with computed tomography. Exp. Biol. San Diego, CA.

Peacham, T.P. and Hedrick, M.S. (2014). Hydrogen sulfide mimics hypoxic respiratory depression in the isolated brainstem of the bullfrog (Lithobates catesbeianus). Exp. Biol. San Diego, CA.

Onyemaechi, C., Crossley II, D.A. and Hedrick, M.S. (2014). Cardiovascular and skeletal muscle sympathetic activity correlates in the embryonic chicken (Gallus domesticus). Exp. Biol. San Diego, CA.

Hedrick, M.S. and McNew, K.A. (2014). Does baroreflex gain vary with habitat in anuran amphibians? Soc. Int. Comp. Biol. Austin, TX.


Grants and Awards

Recent External Awards

NSF      "Collaborative Research: Effects of developmental hypoxia on juvenile cardiac function" $119.167. PI (1/1/18 - 12/31/21). Integrative Organismal Systems. Pending.

NSF      “Experimental tests of a physical model for lymph flow in anurans.” $320,336. Co-PI. (2/15/09-1/31/13). Grant no. IOS 0843082 – Integrative Organismal Systems.

NIH     “Cellular mechanisms for breathing and hypoxia tolerance.” $476,085. PI. (6/1/05-5/31/09). Grant no. SO6 GM48135 (CSUEB MBRS/SCORE program).

NSF      “Developing a comprehensive physical model of lymph movement in amphibians.” $146,344. Co-PI. (8/31/01-9/1/05). Grant no. IBN 0110713 - Integrative Animal Biology.


Professional Service

Editor-in-Chief, Comparative Biochemistry and Physiology, Part A (2013-present)

Editorial Board, Frontiers in Respiratory Physiology (2015-present)

Editorial Board, American Journal of Physiology - Regulatory, Integrative and Comparative Physiology (2013-present)

Editorial Board, Journal of Comparative Physiology B (2012-present)

Editorial Board, Respiratory Physiology and Neurobiology (2008-2012)

Chair, Comparative and Evolutionary Physiology Section of the American Physiological Society (2015-2018)

Service to CSUEB

Chair, Academic Senate (2014-2016)

Chair, Department of Biological Sciences (2008-2011)

Chair, Institutional Animal Care and Use Committee (1998-2008; 2014- )

Other Accomplishments

Awards and Honors

Fellow of the California Academy of Sciences (elected 2008)

George and Miriam Phillips Outstanding Professor Award, CSU, East Bay (2007)

Graduate Students Mentored

CSU, East Bay

Rey Morales (M.S.1997)  

Lise Broch (M.S.2000)

Jennifer Lucitti (M.S. 2001)

Monica Martinez (M.S. 2002)  

Rachel Winmill (M.S. 2003)

Anna Chen (M.S. 2007)

Nick Nimmo (M.S. 2007)

John Polos (M.S. 2007)

Amritha Menon (M.S. 2007)

Mary Ann Maravillas (M.S. 2008)

Eric Wasiolek (M.S. 2008)

Michael Klingler (M.S. 2012)   

University of North Texas

David Oden (M.S. 2013)

Rachel Marschand (M.S. 2013)

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